- Undergraduate student at Faculty of Sciences, Porto University
(1979)
- Graduation in Biology (Licenciatura), Faculty of Sciences, Porto
University (1983)
- MSc in Plant Taxonomy and Ecology, Faculty of Sciences, Porto
University (1989)
- PhD in Plant Taxonomy and Ecology, Faculty of Sciences, Porto
University (1999)
Thesis: "Ecology and Biosystematics of the genus
Sphagnum L. in Portugal"
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